The characters from the animated Madagascar franchise, Alex, Melman, Marty and Gloria celebrate love in this Valentine special.

The Madagascar animals fly back to New York City, but crash-land on an African nature reserve, where they meet others of their own kind, and Alex especially discovers his royal heritage as prince of a lion pride.

The Madagascar animals join a struggling European circus to get back to New York, but find themselves being pursued by a psychotic animal control officer.