The story revolves around sub-zero heroes: a woolly mammoth, a saber-toothed tiger, a sloth and a prehistoric combination of a squirrel and rat, known as Scrat.

When Scrat’s acorn sends an asteroid to Earth, the Herd must find a way to stop the asteroid from hitting Earth with the help of a returning friend.

In the prehistoric past, a young man struggles to return home after being separated from his tribe during a buffalo hunt. He finds a similarly lost wolf companion and starts a friendship that would change humanity.